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Humulus
Family: Cannabaceae
Humulus image
Max Licher
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Ernest Small in Flora of North America (vol. 3)
Herbs , taprooted annuals or rhizomatous perennials, rightward-twining. Stems usually branched, armed with rigid 2-branched, stalked hairs that facilitate climbing. Leaves simple; petioles often twining, with 2-branched hairs. Leaf blade mostly cordate, palmately lobed, sometimes unlobed; surfaces abaxially resin-dotted and×or gland-dotted. Inflorescences: staminate inflorescences axillary and terminal, cymose panicles, erect to pendent, (10-)20-100+-flowered, flowers small; pistillate axillary, spikes or racemes, flowers solitary or paired, short-pedicellate, subtended by bracts and bracteoles. Staminate and pistillate flowers usually on different plants. Achenes lenticular or terete, ensheathed by brownish or sometimes mottled persistent perianth; embryo coiled. x = 10.
JANAS 32(1)
PLANT: Usually twining, dioecious herbs with harsh forked hairs. LEAVES: simple, palmately lobed or unlobed, the abaxial surface often resin dotted. INFLORESCENCE: staminate inflorescences axillary or termi¬nal, 20-100-flowered; pistillate inflorescence axillary. FLOWERS: pistillate flowers paired, subtended by bracts and bracteoles. FRUIT: lenticular, enclosed by persistent perianth. x=10. REFERENCES: Mason, Charles T., Jr. 1999. Cannabaceae. Ariz. - Nev. Acad. Sci. 32(1).
Vascular plants of NE US and adjacent Canada
Dioecious; pistillate fls in short spikes, paired, each pair subtended by a foliaceous bract; ovary enclosed by the membranous, lobeless cal; fr an achene enclosed in the persistent cal and covered by the accrescent bracts; twining, herbaceous, perennial vines with rough stems and broad, opposite, usually lobed lvs. 3, N. Temp.

Gleason, Henry A. & Cronquist, Arthur J. 1991. Manual of vascular plants of northeastern United States and adjacent Canada. lxxv + 910 pp.

©The New York Botanical Garden. All rights reserved. Used by permission.
Species within checklist: Sycamore Canyon, Pajarito Mountains, Santa Cruz
Humulus japonicus
Image of Humulus japonicus
Humulus lupulus
Image of Humulus lupulus
Humulus scandens
Image of Humulus scandens
The National Science Foundation
Development supported by National Science Foundation Grants (DBI 9983132, BRC 0237418, DBI 0743827, DBI 0847966)
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